Maria Liljeroth & Hilary Carlisle 
Ordovician rhynchonelliformean brachiopods from Co. Waterford, SE Ireland 
Palaeobiogeography of the Leinster Terrane

A detailed look at the rhynchonelliformean brachiopods of Ireland This publication, Ordovician rhynchonelliformean brachiopods from Co. Waterford, SE Ireland: Palaeobiogeography of the Leinster Terrane, is Number 62 within the Fossils and Strata series. The international Fossils and Strata series features monographs in palaeontology and biostratigraphy with taxonomic descriptions. The series is owned by and published on behalf of The Lethaia Foundation in cooperation with the Scandinavian countries.

Inhaltsverzeichnis

Introduction 1

Previous work 2

Geological setting 3

Tectonic history of Ganderia and Avalonia 4

The Iapetus Ocean 5

Ireland 6

The Leinster Terrane 7

Docking history 7

Th e Tramore area 7

Material 9

Stratigraphy and fauna 9

Age of the Tramore Limestone Formation 9

Tramore and Dunabrattin/Bunmahon sections 10

Tramore Shale Formation – Tramore area 12

Tramore Limestone Formation – type section at Barrel Strand, Tramore area 12

Lithology 12

Fauna associations 14

Tramore Limestone Formation – other sections 14

Pickardstown sections (Ps) 14

Quillia Section 1 (QS1) 14

Quillia Section 2 (QS2) 14

Bunmahon Formation – Bunmahon/Dunabrattin area 15

Dunabrattin Shale Formation – Bunmahon/Dunabrattin area 15

Dunabrattin Limestone Formation – Bunmahon/ Dunabrattin area 15

Lithology and fauna 16

Correlation 16

Depositional setting and ecology 17

Local events 17

Tramore Limestone Formation 17

Dunabrattin Limestone Formation 19

Palaeobiogeographical reconstructions 20

New reconstructions for the Iapetus Ocean 22

Ganderian terrane positions 22

Longitudinal separation 22

Latitudinal drift of the Ganderian-Avalonian segment 22

Brachiopod provinces 24

Faunal distribution 24

Darriwilian provinces and faunas 26

Th e Low-latitude Province on the Laurentian Platform 26

An early Scoto-Appalachian fauna 27

Th e Toquima-Table Head province 28

Th e mixed faunas of the Leinster Terrane and East Avalonia 31

Th e Leinster Terrane 31

East Avalonia 32

Th e Celtic province 32

Th e Baltic Province 34

Th e High-latitude Province 35

Darriwilian faunal migration patterns: implications on oceanic current confi guration 35

Migrations from the Laurentian margins and intraoceanic islands to the south Iapetus-Tornquist region 38

Migrations within the south Iapetus-Tornquist region 38

Th e Rheic gyre and the High-latitude Province 40

Th e Precordillera 40

Early Sandbian brachiopod provinces 41

Th e Low-latitude Province on the Laurentian Platform 41

Th e Scoto-Appalachian Province 42

Th e Anglo-Welsh-Baltic Province 43

Th e Leinster Terrane 44

East Avalonia and Anglesey 46

Baltica 47

The High-latitude Province 48

The Precordillera 48

Conclusions 48

Systematic palaeontology 50

Abbreviations 50

Repository 50

Statistical methods 50

Allometric growth 50

Phylum Brachiopoda Duméril, 1806 51

Subphylum Rhynchonelliformea Williams, Carlson & Brunton, 1996 51

Class Strophomenata Öpik, 1934 51

Order Strophomenida Öpik, 1934 51

Superfamily Strophomenoidea King, 1846 51

Family Strophomenidae King, 1846 51

Subfamily Strophomeninae King, 1846 51

Genus Tetraphalerella Wang, 1949 51

Tetraphalerella? sp. 52

Subfamily Furcitellinae Williams, 1965 53

Genus Dactylogonia Ulrich & Cooper, 1942 53

Dactylogonia costellata n. sp 53

Family Rafi nesquinidae Schuchert, 1893 55

Subfamily Rafi nesquininae Schuchert, 1893 55

Genus Colaptomena Cooper, 1956 55

Colaptomena auduni n. sp. 56

Colaptomena pseudopecten? (M’Coy, 1846) 57

Superfamily Plectambonitoidea Jones, 1928 59

Family Plectambonitidae Jones, 1928 59

Subfamily Taphrodontinae Cooper, 1956 59

Genus Isophragma Cooper, 1956 59

Isophragma parallelum n. sp. 59

Family Leptellinidae Ulrich & Cooper, 1936 61

Subfamily Leptellininae Ulrich & Cooper, 1936 61

Genus Leptellina Ulrich & Cooper, 1936 61

Leptellina llandeiloensis (Davidson, 1883) 61

Subfamily Palaeostrophomeninae Cocks & Rong, 1989 62

Genus Glyptambonites Cooper, 1956 62

Glyptambonites sp. 62

Family Grorudiidae Cocks & Rong, 1989 63

Genus Grorudia Spjeldnæs, 1957 63

Grorudia grorudi Spjeldnæs, 1957 63

Family Leptestiidae Öpik, 1933a, emend. Cocks & Rong, 1989 64

Genus Leptestiina Öpik, 1933a 64

Leptestiina derfelensis (Jones, 1928) 65

Family Sowerbyellidae Öpik, 1930 66

Subfamily Sowerbyellinae Öpik, 1930 66

Genus Sowerbyella Jones, 1928 66

Subgenus Sowerbyella (Sowerbyella) Jones, 1928 66

Sowerbyella (Sowerbyella) antiqua Jones, 1928 66

Order Billingsellida Schuchert, 1893 70

Suborder Clitambonitidina Öpik, 1934 70

Superfamily Clitambonitoidea Winchell & Schuchert, 1893 70

Family Clitambonitidae Winchell & Schuchert, 1893 70

Genus Atelelasma Cooper, 1956 70

Atelelasma longisulcum n. sp. 70

Class Rhynchonellata Williams, Carlson & Brunton, 1996 73

Order Orthida Schuchert & Cooper, 1932 73

Suborder Orthidina Schuchert & Cooper, 1932 73

Superfamily Orthoidea Woodward, 1852 73

Family Orthidae Woodward, 1852 73

Genus Sulevorthis Jaanusson & Bassettt, 1993 73

Sulevorthis aff S. blountensis (Cooper, 1956) 73

Family Glyptorthidae Schuchert and Cooper, 1931 74

Genus Glyptorthis Foerste, 1914 74

Glyptorthis crispa (M’Coy, 1846) 74

Family Hesperorthidae Schuchert & Cooper, 1931 77

Genus Hesperorthis Schuchert & Cooper, 1931 77

Hesperorthis leinsterensis n. sp. 77

Family Plaesiomyidae Schuchert, 1913 86

Subfamily Plaesiomyinae Schuchert, 1913 86

Genus Valcourea Raymond, 1911 86

Valcourea confinis (Salter, 1849) 86

Family Productorthidae Schuchert & Cooper, 1931 87

Subfamily Productorthinae Schuchert & Cooper, 1931 87

Genus Productorthis Koz?owski, 1927 87

Productorthis sp 87

Superfamily Plectorthoidea Schuchert & Le Vene, 1929 89

Family Platystrophiidae Schuchert & Le Vene, 1929, emend.

Zuykov & Harper, 2007 89

Genus Platystrophia King, 1850, emend. Zuykov & Harper, 2007 89

Platystrophia tramorensis n. sp. 89

Platystrophia aff P. sublimis Öpik, 1930 92

Suborder Dalmanellidina Moore, 1952 94

Superfamily Dalmanelloidea Schuchert, 1913 94

Family Dalmanellidae Schuchert, 1913 94

Subfamily Dalmanellinae Schuchert, 1913 94

Genus Howellites Bancroft , 1945 94

Howellites hibernicus n. sp. 94

Family Paurorthidae Öpik, 1933b 106

Genus Paurorthis Schuchert & Cooper, 1931 106

Paurorthis aff P. parva (Pander, 1830) 106

Superfamily Enteletoidea Waagen, 1884 110

Family Linoporellidae Schuchert & Cooper, 1931 110

Genus Salopia Williams in Whittington & Williams, 1955 110

Salopia gracilis Williams in Whittington & Williams, 1955 110

Order Pentamerida Schuchert & Cooper, 1931 111

Suborder Syntrophiidina Ulrich & Cooper, 1936 111

Superfamily Syntrophioidea Ulrich & Cooper, 1936 111

Family Porambonitidae Ulrich & Cooper, 1936 111

Genus Hibernobonites n. gen. 111

Hibernobonites filosus (M’Coy, 1846) 112

Acknowledgements 115

References 115

Plates 1-10 122

Appendix A 142

Appendix B 148

Über den Autor

Maria Liljeroth is the author of Ordovician rhynchonelliformean brachiopods from Co. Waterford, SE Ireland: Palaeobiogeography of the Leinster Terrane, published by Wiley. David A.T. Harper is a British palaeontologist, specialising in fossil brachiopods and numerical methods in palaeontology. He is Professor of Palaeontology in Earth Sciences, Principal of Van Mildert College, and Deputy Head of Colleges in Durham University.